Plant traitPreferred unitRange of valuesRecommended no. of replicates 
   MinimumPreferredCV Range (%)
2 Whole-plant traits
2.1 Life historyCategorical35
2.2 Life formCategorical35
2.3 Growth formCategorical35
2.4 Plant heightm< 0.01–140102517–36
2.5 ClonalityCategorical510
2.6 Spinescence510
Spine lengthmm0.5–300
Spine widthmm0.5–30
Spine length: leaf lengthUnitless0–30
2.7 Branching architectureNo. of ramifications per branch0 – > 100510
2.8 Leaf to sapwood areaUnitless100–103510
2.9 Root-mass fractionUnitless0.15–0.40
(for seedlings, down to 0.10)
510
2.10 Salt-tolerance traits510
Selective root cation uptakeUnitless10
Salt excretion and compartmentalisationCategorical5
2.11 Relative growth ratemg g-1 day-12–3001020
2.12 Shoot flammabilityUnitless0 – ~3510
2.13 Water-flux traits1020
Gap fractionUnitless0–1
Stem flow%0–50
Water retention on plant surfaceg m-20–500
Leaf wettabilitydegrees (contact angle)0–180
Droplet retention abilitydegrees (slope angle)0–90
3  Leaf traits
3.1  Specific leaf aream2 kg-1 (mm2 mg-1)A< 1–3005, 510, 48–16
3.2 Area of a leafmm21 – > 2065, 510, 417–36
3.3 Leaf dry-matter contentmg g-150–7005, 510, 44–10
3.4 Leaf thicknessmm< 0.1–5B510
3.5 pH of green leaves or leaf litterUnitless3.5–6.55, 510, 41–6
3.6 Leaf nitrogen and phosphorus concentrations
Leaf nitrogen concentrationmg g-15–705, 510, 48–19
Leaf phosphorus concentrationmg g-10.2–55, 510, 410–28
3.7 Physical strength of leaves5, 510, 414–29
Force to tearN mm-10.17–40
Work to shearJ m-10.02–0.5
Force to punchN mm-10.03–1.6
3.8 Leaf lifespan and duration of green foliage
Leaf lifespanMonth0.5–2005, 4010, 16011–39
Duration of green foliageMonth1–12510
3.9 Vein densityMm mm-20.5–25510
3.10 Light-saturated photosynthetic rateµmol m–2 s–12–30510
3.11 Leaf dark respirationµmol m–2 s–10.4–4510
3.12 Photosynthetic pathwayCategorical33
3.13 C-isotope composition–35 – –10510
3.14 Electrolyte leakage%2–1005, 510, 49–26
3.15 Leaf water potentialMPa–7 —> 05, 55, 1011–33
3.16 Leaf palatability% Leaf area consumed0–1001020
3.17 Litter decomposabilityC% Mass loss0–10010207–14
4 Stem traits
4.1 Stem-specific densitymg mm–3 (kg l–1)A0.1–1.35105–9
4.2 Twig dry-matter contentmg g-1150–8505102–8
4.3 Bark thicknessmm0.1 – > 30510
4.4 Xylem conductivity51021–63
Stem-specific xylem hydraulic conductivity (KS)kgm–1 s–1 MPa–11 (gymnosperms) to 200 (tropical lianas)
Leaf-area-specific xylem hydraulic conductivity (KL)kgm–1 s–1 MPa–16 X 10-5 (gymnosperms) to 1 X 10-2 (tropical lianas)
4.5 Vulnerability to embolism (y50)MPa–0.25–1451020–45
5 Below-ground traits
5.1 Specific root length                                                                                                                                         m g-1       3–350    5, 1010, 10       15–24
5.2 Root-system morphology510
Depthm0.05–70
Lateral extentm0.05–40
Density of explorationmm mm–310-4–1
5.3 Nutrient uptake strategyCategorical510
6 Regenerative traits
6.1 Dispersal syndrome
Categorical36
6.2 Dispersule size and shape
Size (mass)mg (g)A510
ShapeUnitless0–136
6.3 Dispersal potentialDispersules1020
6.4  Seed massmg10-3–10751014–27
6.5  Seedling morphologyCategorical36
6.6  Resprouting capacityUnitless0–100510